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From this viewpoint, the effect of the electron acceptor condition is one of the most important
questions in examining actual EBPR processes, especially in regard to nitrogen removal. Kerrn-
Jespersen and Henze (1993) suggested that a part of PAOs (denitrifying-PAOs) could utilize both
oxygen and nitrate as electron acceptor while the others could utilize oxygen only
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"The coral itself has more energy for growth because the reduced zooxanthellae density requires less energy."
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